3Graduate School of Engineering, Kobe University, 1-1 Rokkodai, Nada-ku, Kobe 657-8501, Japan

†Present address: KYOUSEI Science Center for Life and Nature, Nara Women’s University, Kitauoyahigashimachi, Nara, Nara 630 8506, Japan.

‡Present address: Morphogenetic Signaling Team, RIKEN Center for Biosystems Dynamics Research, Chuo-ku, Kobe 650-0047, Japan.

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